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1 Introduction Plants rely on intricate signal perception and regulatory mechanisms to respond to internal and external stimuli during growth, development, and adaptation to environmental stresses ( The WAK gene family was initially identified in Arabidopsis ( SlWAK1 gene impairs salt tolerance throughout the plants life cycle, primarily due to its failure to maintain osmotic homeostasis ( GhWAKL26 may positively regulate salt tolerance by modulating the expression of ion transport-related genes, thereby altering the intracellular levels of Na + and K + ( WAK genes have been extensively studied in various plants, such as rice ( N

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